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Books Received
Primate-Science / PrimateLit


MANAGEMENT OF LORISES IN CAPTIVITY
A HUSBANDRY MANUAL FOR ASIAN LORISINES (NYCTICEBUS & LORIS SSP.)

Edited by: Helena Fitch-Snyder and Helga Schulze


Compiler: Lena C. Larson
Illustrations: Helga Schulze



Published by the Center for Reproduction of Endangered Species (CRES)
Zoological Society of San Diego 2001



INTRODUCTION


The purpose of this manual is to provide basic husbandry guidelines to loris managers, caretakers, and veterinarians.
The information in this manual has been compiled from numerous articles, reports, and personal experiences by the
authors and editors. Because new information is continuously being discovered, it is tempting to keep delaying
publication until the husbandry manual is even more complete. However, as an increasing number of institutions become
involved with the management of these specialized prosimians, it has become essential to have this information 
vailable in a usable format.


When the pygmy loris was first introduced to the Western Hemisphere a little over a decade ago, its basic biology
and husbandry requirements were virtually unknown. Pygmy loris management techniques were derived from methods that
had previously been developed for slow and slender lorises. Researchers and managers are just beginning to understand
some of the characteristics that make the pygmy loris unique.


The three lords types each have unique characteristics that are species specific. However, because they share many
of the same management methods, housing conditions, and behavioral characteristics, the editors decided that information
about all the three loris species should be included in this volume.


In 1995, The Pygmy Loris Species Survival Plan (SSP) was established through the American Association of Zoos and
Aquariums (AZA). One of the goals of this group is to develop a self-sustaining captive population of this species.
Along with the Pygmy Loris Masterplan (Fitch-Snyder, 1998), this husbandry manual is intended as a tool to assist
toward this goal.


The editors plan to periodically distribute updated and new information to SSP Institution Representatives and other
appropriate recipients. Users of this publication are encouraged to maintain this manual as a working document and
contribute additional material for future updates.


Helga Schulze is compiling additional information, particularly for slender loris husbandry (see also Schulze, 1998);
readers who are interested can contact her directly via e-mail (helga.schulze@cityweb.de) or at her University address,
which is listed in the authors section of this publication.


In addition to printed materials, some loris information is available via the Internet. Here, new data and amendments
to printed information may be published in the future. Two addresses, as an example: Loris homepage by Dr. K. I. Anna
Nekaris, Assistant Professor, Department of Anthropology, Southern Illinois University;
address: http://www.nocturnalprimate.org. In http://www.species.net, a web site of Quantum Conservation for husbandry
and conservation information (by Richard Perron), a loris conservation database is under construction (compiler: Helga
Schulze; direct address: http:// www.species.net).


Helena Fitch-Snyder



CONTENTS


Taxonomy and Description 1
Distribution an Status 7
History of Lorises in Captivity 13
Behavior 14
Introduction 14
Social Behavior 4
Housing and Social Behavior 15
Aggressive Behavior 16
Loris Ethogram 18
Reproduction 28
Determining Gender 28
Adolescent Development 29
Estrous Cycles 30
Gonadal Morphology in Males 30
Copulation 32
Gestation 33
Parturition 33
Reaction of Others to Newborns 34
Resumption of Estrus 35
Interbirth Interval 35
Reproductive Seasonality 36
Litter Size 38
Infant Development and Parental Behavior 39
Captive Reproductive History 41
Infant Care 45
Birth Management and Hand Rearing
-contributed by Janet Hawes and Jacqueline Ogden 45
Hand Rearing 45
Post-Partum Observations and Evaluation 45
Alternatives to Hand Rearing 45
Physical Needs 46
Physical Needs: Species-Specific Concerns 49
Social Needs 50
Diet 52
Diet Composition in the Wild 52
Diet Composition in Captivity 53
Diet Composition in Captivity for the Pygmy Loris
-contributed by Barbara Toddes and Kerri Slifka 55
Health 60
Review of Loris Clinical Information and
Pathology Data from the San Diego Zoo: 1982- 1995
-contributed by Meg Sutherland Smith, D.V.M. and Ilse Stalis, D.V.M. 60
Introduction 60
Preventive Medicine 60
Anesthesia 60
Clinical Pathology 61
Medical Review 61
Summary 65
Tables 66
Habitat Design 71
Minimum Standards for Housing Asian Lorisines
(Adapted from AZA guidelines for the Family Lorisidae)
-contributed by Lisa Bottcher-Law 71
General Habitat Design 72
Temperature Levels 78
Lighting Conditions 78
Cleaning Cages 79
An Enriched Environment
--contributed by Lisa Bottcher-Law 80
Mixed Species Housing
-contributed by Barbara Lester 88
Literature Cited 93
Map references 100
N. coucang distribution map (Figure 9) 100
N. pygmaeus distribution map (Figure 10) 101
Loris distribution maps (Figures 11 & 12) 102



HOW TO OBTAIN:


To borrow this document, contact Joanne Brown (brown@primate.wisc.edu), Head of Technical Services, Wisconsin
Primate Research Center Library and Information Service, University of Wisconsin-Madison. Fax: 1-608-263-3512


While preference should be give to borrowing the WPRC copy, Helena indicated she would try to accommodate people
who have a significant need for their own copy. Write her at:


Fitch-Snyder, Helena
nycticebus@yahoo.com

URL: http://www.primate.wisc.edu/pin/review/management.html
Page last modified: November 1, 2002
Maintained by the WRPRC Library

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