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SPECIES AND SUBSPECIES OF PRIMATES DESCRIBED SINCE 1990

By Anthony Rylands
a.rylands@conservation.org

Thirty-eight primates, species and subspecies, have been described in the last ten years: 10 from Madagascar, eight from Africa, seven from South-east Asia, 10 from the Brazilian Amazon (seven of them marmosets), and three from the Brazilian Atlantic forest. Please note that this list is presented without any judgement as to the validity or otherwise of the various primates described.

Prosimians

1. Pseudopotto martini Schwartz, 1996 (False potto)
2. Tarsius dianae Niemitz, Nietsch, Warter & Rumpler 1991 (Dian's tarsier)
3. Galagoides rondoensis Honess 1996 (Rondo dwarf galago)*
4. Galagoides udzungwensis Honess 1996 (Matundu dwarf galago)*
5. Microcebus ravelobensis Zimmerman, Cepok, Rakotoarison, Zietemann & Radespiel, 1998 (Lac Ravelobe or golden-brown mouse lemur)
6. Microcebus tavaratra Rasoloarison, Goodman & Ganzhorn, 2000 (Northern rufous mouse lemur)
7. Microcebus sambiranensis Rasoloarison, Goodman & Ganzhorn, 2000 (Sambirano mouse lemur)
8. Microcebus berthae Rasoloarison, Goodman & Ganzhorn, 2000 (Berthe's mouse lemur)
9. Cheirogaleus ravus Groves, 2000 (Large iron-grey dwarf lemur)
10. Cheirogaleus minusculus Groves, 2000 (Lesser iron-grey dwarf lemur)
11. Phaner furcifer pallescens Groves & Tattersall, 1991 (Western fork-crowned lemur)
12. Phaner furcifer parienti Groves & Tattersall, 1991 (Sambirano fork-crowned lemur)
13. Phaner furcifer electromontis Groves & Tattersall, 1991 (Amber mountain fork-crowned lemur)
14. Avahi unicolor Thalmann & Geissmann, 2000 (Unicolor avahi)

* The first descriptions were in "Honess, P. E. 1996. Speciation Among Galagos (Primates: Galagidae) in Tanzanian Forests. Ph.D thesis, Oxford Brookes University, Oxford, UK". While here attributed to Honess (1996), both forms were described and illustrated in Kingdon (1997). Not accepting the validity of a doctoral thesis as a formal published description, the authorship is also attributed to "Honess, 1997" by Groves (2001), referring to the descriptions in Kingdon (1997). Honess and Bearder (1996) also published the descriptions of these species, but issue 2(2), 1996, of African Primates was published after Kingdon (1997).

Old World monkeys

15. Miopithecus ogouensis Kingdon, 1997 (Northern talapoin)
16. Cercopithecus cephus ngottoensis Colyn, 1999 (Ngotto moustached monkey)
17. Cercopithecus erythrogaster pococki Grubb, Lernould & Oates, 1999 (Nigerian white-throated guenon)
18. Macaca pagensis siberu Fuentes & Olson, 1995 (Siberut macaque)
19. Colobus badius semlikiensis Colyn, 1991
20. Procolobus badius epieni Grubb & Powell, 1999 (Niger Delta red colobus)
21. Presbytis melalophos bicolor Aimi and Bakar, 1992 (Sumatran sureli)
22. Semnopithecus auratus ebenus Brandon-Jones, 1995 (Wulsin's ebony leaf monkey)
23. Pygathrix nemaeus cinerea Nadler, 1997 (Grey-shanked douc langur)
24. Rhinopithecus roxellana hubeiensis Wang, Jiang & Li, 1998 (Hubei golden snub-nosed monkey)
25. Rhinopithecus roxellana qinlingensis Wang, Jiang & Li, 1998 (Qinling golden snub-nosed monkey)

New World monkeys

26. Callithrix nigriceps Ferrari and Lopes, 1992 (Black-headed marmoset)
27. Callithrix mauesi Mittermeier, Ayres & Schwarz, 1992 (Maues marmoset)
28. Callithrix argentata marcai Alperin, 1993 (Marca's marmoset)
29. Callithrix saterei Souse e Silva Jr & Noronha, 1998 (Satere marmoset)
30. Callithrix humilis Van Roosmalen, Van Roosmalen, Mittermeier & Fonseca, 1998 (Black-crowned dwarf marmoset)
31. Callithrix manicorensis Van Roosmalen, Van Roosmalen, Mittermeier & Rylands, 2000 (Manicore marmoset)
32. Callithrix acariensis (Van Roosmalen, Van Roosmalen, Mittermeier & Rylands, 2000 (Rio Acari marmoset)
33. Leontopithecus caissara Lorini and Persson, 1990 (Black-faced lion tamarin)
34. Callicebus personatus barbarabrownae Hershkovitz, 1990 (Blond titi)
35. Callicebus coimbrai Kobayashi & Langguth 1999 (Coimbra-Filho's titi monkey)
36. Cebus kaapori Queiroz, 1992 (Ka'apor capuchin)
37. Callicebus bernardi, 2002 (Bernardi monkey)
38. Callicebus stephennashi, 2002

References

Aimi, M. and Bakar, A. 1992. Taxonomy and distribution of Presbytis melalophos group in Sumatra, Indonesia. Primates 33: 191-206.
Alperin, R. 1993. Callithrix argentata (Linnaeus, 1771): considera‡oes taxon“micas e descri‡ao de subesp‚cie nova. Bol. Mus. Para. Em¡lio Goeldi, S‚r. Zool. 9(2): 317- 328.
Brandon-Jones, D. 1995. A revision of the Asian pied leaf monkeys (Mammalia: Cercopithecidae; Superspecies Semnopithecus auratus), with a description of a new subspecies. Raffles Bulletin of Zoology 43(1): 3-43.
Colyn, M. 1991. L' importance zoog‚ographique du bassin du fleuve Za‹re pour la speciation: La cas des Primates Simiens. Koninklijk Museum voor Midden-Afrika, Tervuren, Belgi‰, Annalen. Zoologische Wetenschappen 264: i-ix, 1-250.
Colyn, M. 1999. Etude populationnelle de la super-espŠce Cercopithecus cephus habitant l'enclave forestiŠre Sangha-Oubangui (R‚publique Centrafricaine). Description de C. cephus ngottoensis subsp.nov.) Mammalia 63(2): 137-147.
Ferrari, S. F. and Lopes, M.A. 1992. A new species of marmoset, genus Callithrix Erxleben 1777 (Callitrichidae, Primates) from western Brazilian Amazonia. Goeldiana Zoologia (12): 1-3.
Fuentes, A. and Olson, M. 1995. Preliminary observations and status of the Pagai macaque. Asian Primates 4(4): 1-4. (See Groves, C. P. 1986. The nomenclature of the Tanzanian mangabey and the Siberut macaque. Australian Primatol. 10(4): 2-5.) (See also: Whitten, A. J. and Whitten, J. E. J. 1982. Preliminary observations of the Mentawai macaque on Siberut Island, Indonesia. Int. J. Primatol. 3: 445-459)
Groves, C. P. 2000. The genus Cheirogaleus: Unrecognized biodiversity in dwarf lemurs. Int. J. Primatol. 21(6): 943-962.
Groves, C. P. 2001. Primate Taxonomy. Smithsonian Institution Press, Washington, DC.
Grubb, P. and Powell, C. B. 1999. Discovery of red colobus monkeys (Procolobus badius) in the Niger Delta with the description of a new and geographically isolated subspecies. J. Zool., Lond. 248: 67-73.
Grubb, P., Lernould, J.-M. and Oates, J. F. 1999. Validation of Cercopithecus eythrogaster pococki as the name for the Nigerian white-throated guenon. Mammalia 63(3): 389-392.
Hershkovitz, P. 1990. Titis, New World monkeys of the genus Callicebus (Cebidae, Platyrrhini): a preliminary taxonomic review. Fieldiana, Zoology, New Series, (55): 1-109.
Honess, P. E. 1996. Speciation Among Galagos (Primates: Galagidae) in Tanzanian Forests. Ph.D thesis, Oxford Brookes University, Oxford, UK Honess, P. E. and Bearder, S. 1996. Descriptions of the dwarf galago species of Tanzania. African Primates 2(2); 75-79.
Kingdon, J. 1997. The Kingdon Field Guide to African Mammals. Academic Press, San Diego.
Kobayashi, S. and Langguth, A. 1999. A new species of titi monkey, Callicebus Thomas, from north-eastern Brazil (Primates, Cebidae). Revta. Bras. Zool. 16(2): 531-551.
Mittermeier, R. A., Schwarz, M. and Ayres, J. M. 1992a. A new species of marmoset, genus Callithrix Erxleben 1777 (Callitrichidae, Primates), from the Rio Mau‚s region, state of Amazonas, Central Brazilian Amazonia. Goeldiana Zoologia (14): 1-17.
Nadler, T. von. 1997. A new subspecies of douc langur, Pygathrix nemaeus cinereus ssp. nov. Zool. Garten N. Z. 67(4): 165-176.
Niemitz, C., Nietsch, A., Warter, S. and Rumpler Y. 1991. Tarsius dianae: a new primate species from Central Sulawesi (Indonesia). Folia Primatol. 56:105-116.
Queiroz, H. L. 1992. A new species of capuchin monkey, genus Cebus Erxleben 1977 (Cebidae, Primates), from eastern Brazilian Amazonia. Goeldiana Zoologia (15): 1-3.
Rasoloarison, R. M., Goodman, S. M. and Ganzhorn, J. U. 2000. Taxonomic revision of mouse lemurs (Microcebus) in the western portions of Madagascar. Int. J. Primatol. 21(6): 963-1019.
Schwartz, J., H. 1996. Pseudopotto martini: A new genus and species of extant lorisiforme primate. Anthropological Papers of the American Museum of Natural History 78: 1-14. (See also Groves, C. P. 1997-1998. Pseudopotto martini: A new potto? African Primates 3(1-2): 42-43; Bearder, S. 1997-1998. Pseudopotto: When is a potto not a potto? African Primates 3(1-2): 43-44; Sarmiento, E. 1997-1998. The validity of "Pseudopotto martini". African Primates 3(1-2): 44-45.)
Silva Jr., J. S. and Noronha, M. de A. 1998. On a new species of bare-eared marmoset, genus Callithrix Erxleben, 1777, from central Amazonia, Brazil (Primates: Callitrichidae). Goeldiana Zoologia (21): 1-28.
Thalmann, U. and Geissmann, T. 2000. Distributions and geographic variation in the western woolly lemur (Avahi occidentalis) with description of a new species (A. unicolor). Int. J. Primatol. 21(6): 915-941.
Van Roosmalen, M. G. M., Van Roosmalen, T., Mittermeier, R. A. and Fonseca, G. A. B. da. 1998. A new and distinctive species of marmoset (Callitrichidae, Primates) from the lower Rio Aripuana, state of Amazonas, central Brazilian Amazonia. Goeldiana Zoologia (22): 1-27.
Van Roosmalen, M. G. M., Van Roosmalen, T., Mittermeier, R. A. and Rylands, A. B. 2000. Two new Species of marmoset, genus Callithrix Erxleben, 1777 (Callitrichidae, Primates), from the Tapaj¢s/Madeira interfluvium, south central Amazonia, Brazil. Neotrop. Primates 8(1): 1-18.
Wang Y., Jiang X. and Lid D. 1998. Classification and distribution of the extant subspecies of golden snub-nosed monkey (Rhinopithecus [Rhinopithecus] roxellana). In: The Natural History of the Doucs and Snub-nosed Monkeys, N. G. Jablonski (ed.), pp.53-64. Recent Advances in Human Biology, Vol. 4. World Scientific Publishing Company, Singapore. (See also: Wang Y., Jiang X. and Lid D. 1994. Classification of existing subspecies on golden snub-nosed monkey, Rhinopithecus roxellana (Colobinae, Primates). In: Handbook and Absrtracts. XVth Congress of the International Primatological Society, 3-8 August 1994, Kuta, Bali, Indonesia, p.277. Abstract)
Zimmerman, E., Cepok, S., Rakotoarison, N., Zietemann, V. and Radespiel, U. 1998. Sympatric mouse lemurs in north-west Madagascar: A new rufous mouse lemur species. Folia Primatol. 69: 106-114.

Anthony B. Rylands
Deputy Chair - IUCN/SSC Primate Specialist Group
Center for Applied Biodiversity Science
Conservation International
1919 M. Street NW
Suite 600
Washington, DC 20036
USA
Tel: (202) 912 1714
E-mail: a.rylands@conservation.org


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